Arrested for reckless space driving, super human Chiharu and cyborg Makina are sentenced to clean the Milky☆Subway—an old interplanetary train. Easy enough, right? Wrong. When the train suddenly takes off, they are thrown into a high-speed space-train spectacle packed with chaos, no plans, and zero strategy. Just pure momentum!

After graduating from high school, Natsuko Hirose starts her career as an animator. Her talent quickly flourishes, and she makes her debut as a director in no time. Her first anime becomes a massive hit, sparking a social phenomenon and earning her recognition as an up-and-coming genius director. Her next project is set to be a romantic comedy movie themed around first love! However, having never been in love herself, Natsuko struggles to understand the concept of first love, and as a result, she’s unable to create the storyboard, causing the movie production to come to a standstill. One day, she passes out while working on her storyboard and wakes up in the world of her favorite childhood anime movie "A Tale of Perishing."