A terminally ill girl named Himari Takakura is miraculously saved from death by a strange spirit who resides in a penguin-shaped hat. However, in exchange for extending her life, the spirit tasks Himari's brothers, Kanba and Shōma, to seek out an elusive item known as the Penguin Drum with assistance from a trio of strange penguins.

Makoto 'Smile' Tsukimoto and his friend Yutaka 'Peco' Hoshino have been playing table tennis together since they were kids, but as they enter high school, they find that the game, and how they see it, has changed. Peco, brimming with confidence and energy, wants to be the best in the world, but the reserved Smile has little fighting spirit and doesn't want to sacrifice others' happiness just to win, despite his innate talent. As the two grow, and experience the ups and downs of the sport – and life – they try to figure out exactly who they really are and what drives them to play.

With a demonic, soul-consuming weapon in hand, can a legendary swordsman and a samurai brotherhood defeat a bloodthirsty zombie uprising?